Oppenheim, 1930

Dungulia Oppenheim, 1930 is a junior synonym (available and invalid), see Smilotrochus Milne Edwards and Haime, 1851, p. 58.

Type Species

Coelosmilia milneri Gregory, 1898, p. 249.; Original Designation Gregory, 1898, p. 249.

Type Specimen: Holotype; ; Not Traced; Unknown

Type Locality: Lower Eocene of Egypt (Dungul Wells).



Gregory (1898) described the solitary species Coelosmilia milneri which, according to Gregory (1898, p. 249), is distinguished from the genus Smilotrochus only by “less simple and less crowded septa”, as well as “broader interseptal loculi”. While it remains unclear what exactly Gregory referred to as “simple” or “less simple” septa, respectively, the character of density of septa (Gregory used the term “crowded septa”, resulting in “narrower interseptal loculi”) could be considered a specific but not a generic feature. Therefore, the species Coelosmilia milneri (type species of the genus Dungulia) is considered a species of Smilotrochus. Hence, because it is the type species of Dungulia, the latter represents a junior synonym of Smilotrochus (Baron-Szabo, 2008). In creating the genus Dungulia, Oppenheim (1930b) used non-type material to prove and establish a relation to the dendrophylliid group of eupsammiids. Because in the original description of Coelosmilia milneri, Gregory (1898) solely compared his new species with other caryophylliid species, a relation only with caryophylliid forms can be assumed. However, the presence of Trochosmilia-like septa (=rather thin, often finely granulated septa) which have the tendency to fuse was already documented by Milne Edwards and Haime (1850) and later included in the first generic description of Smilotrochus by Milne Edwards (1857, vol. II, p. 70). Especially in taxa of this genus which have flexuous septa, as already reported for the Maastrichtian form Smilotrochus hagenowi Milne Edwards (1857, vol. 2, p. 71), the fusion of their axial ends can result in a septal pattern that resembles the dendrophyllid type (compare specimens figured in Baron-Szabo, 2008, Pl. 7, figs 5a and 6–8).


  • North Africa; Eocene

This page has been in preparation since 02-Jun-2007 12:35

This version was contributed by Rosemarie Baron-Szabo on 14-Sep-2010 21:22.

Page authors are: Rosemarie Baron-Szabo. Please contact the editor if you would like to contribute to the diagnosis of this taxon.

The editor is: Rosemarie Baron-Szabo

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